:root {
  color-scheme: light;
  --ink: #202625;
  --muted: #65706c;
  --accent: #eecb3b;
  --line: #dfe3df;
  --paper: #fff;
  --wash: #f5f6f3;
  --width: 1120px;
}
* { box-sizing: border-box; }
html { scroll-behavior: smooth; scroll-padding-top: 96px; }
body { margin: 0; background: var(--paper); color: var(--ink); font-family: "Helvetica Neue", Helvetica, Arial, sans-serif; font-size: 17px; line-height: 1.7; -webkit-font-smoothing: antialiased; }
[hidden] { display: none !important; }
a { color: inherit; text-underline-offset: .22em; }
button { font: inherit; }
a, button, summary { -webkit-tap-highlight-color: transparent; }
a:focus-visible, button:focus-visible, summary:focus-visible, [tabindex]:focus-visible { outline: 3px solid #a57e00; outline-offset: 5px; border-radius: 2px; }
.hero a:focus-visible, .hero button:focus-visible { outline-color: var(--accent); }
svg { width: 22px; height: 22px; flex-shrink: 0; }
p { margin: 0 0 1.25rem; }
h1, h2, h3 { margin: 0; font-weight: 500; letter-spacing: -.045em; }
h2 { font-size: clamp(2.25rem, 4.2vw, 3.55rem); line-height: 1.13; }
h3 { font-size: 2rem; line-height: 1.22; }
.container { width: min(var(--width), calc(100% - 96px)); margin-inline: auto; }
.eyebrow { margin: 0 0 1.4rem; font-size: .72rem; font-weight: 600; line-height: 1.5; letter-spacing: .17em; text-transform: uppercase; }
.skip-link { position: fixed; z-index: 20; top: -100px; left: 16px; padding: 10px 20px; background: white; color: var(--ink); }
.skip-link:focus { top: 16px; }
.hero { position: relative; min-height: max(800px, 100svh); isolation: isolate; color: white; background: #222624; overflow: hidden; }
.hero-media, .hero-shade { position: absolute; inset: 0; z-index: -1; }
.hero-media {
  transform: translateX(30%);
  -webkit-mask-image: linear-gradient(to right, transparent, #000 12%);
  mask-image: linear-gradient(to right, transparent, #000 12%);
}
.hero-media img, .hero-media video { position: absolute; inset: 0; width: 100%; height: 100%; object-fit: cover; object-position: 66% center; }
.hero-media video { opacity: 0; transition: opacity .4s ease; }
.hero-media video.is-playing { opacity: 1; }
.hero-shade { background: linear-gradient(90deg, rgba(13,19,18,.9) 0%, rgba(13,19,18,.74) 30%, rgba(13,19,18,.19) 75%, rgba(13,19,18,.08)), linear-gradient(0deg, rgba(13,19,18,.52), transparent 25%, transparent 82%, rgba(13,19,18,.45)); }
.hero-nav, .hero-content, .hero-bottom { width: min(1360px, calc(100% - 112px)); margin-inline: auto; }
.hero-nav { display: flex; align-items: center; justify-content: space-between; padding-top: 36px; gap: 24px; }
.wordmark { display: inline-flex; align-items: center; gap: 11px; text-decoration: none; }
.wordmark svg { width: 35px; height: 35px; color: var(--accent); }
.wordmark span { font-size: 10px; font-weight: 600; line-height: 1.35; letter-spacing: .16em; }
.hero-nav-links { display: flex; gap: 34px; font-size: .8rem; }
.hero-nav-links a { text-decoration: none; padding: 8px 0; color: #e8ece9; }
.hero-nav-links a:hover { color: var(--accent); }
.hero-content { padding-top: clamp(74px, 10vh, 130px); padding-bottom: 130px; }
.hero-eyebrow { color: #e0e3d9; display: flex; align-items: center; gap: 10px; font-size: .66rem; }
.hero-eyebrow span { width: 6px; height: 6px; border-radius: 50%; background: var(--accent); }
h1 { font-size: clamp(4.5rem, 7.1vw, 7rem); line-height: 1.01; letter-spacing: -.055em; }
h1 span { color: var(--accent); }
.hero-summary { margin-top: 26px; color: #e0e4df; font-size: 1rem; line-height: 1.65; max-width: 440px; }
.hero-actions { display: flex; gap: 12px; flex-wrap: wrap; margin-top: 28px; }
.button { display: inline-flex; align-items: center; justify-content: center; min-height: 48px; gap: 20px; padding: 11px 19px; font-size: .78rem; font-weight: 500; text-decoration: none; border: 1px solid transparent; border-radius: 4px; transition: background .2s, border-color .2s; }
.button svg { width: 18px; height: 18px; }
.button-primary { background: var(--accent); color: #242818; }
.button-primary:hover { background: #ffdf64; }
.button-outline { border-color: #9baba077; color: #fff; background: #17201b33; }
.button-outline:hover { background: #ffffff17; border-color: #fff8; }
.button-note { font-size: .6rem; letter-spacing: .05em; color: #d6ded8; }
.paper-authors { max-width: 660px; margin-top: 25px; font-size: .9rem; }
.author-list { display: flex; flex-wrap: wrap; gap: 8px 22px; margin: 0; color: #fff; }
.author-list > span { white-space: nowrap; }
.author-list a { text-decoration-color: #ffffff60; text-decoration-thickness: 1px; }
.author-list a:hover { color: var(--accent); text-decoration-color: currentColor; }
.paper-authors sup { color: var(--accent); font-size: .67em; line-height: 0; margin-left: 3px; }
.author-affiliations { list-style: none; padding: 0; margin: 14px 0 12px; color: #e0e4df; font-size: .74rem; line-height: 1.8; }
.author-affiliations li { padding-left: 14px; position: relative; }
.author-affiliations sup { position: absolute; left: 0; top: .9em; margin: 0; font-size: .78em; }
.author-notes { display: grid; gap: 2px; margin: 0; color: #c8d0c8; font-size: .68rem; line-height: 1.7; }
.author-notes sup { margin: 0 4px 0 0; font-size: .9em; }
.hero-bottom { position: absolute; bottom: 32px; inset-inline: 0; display: flex; justify-content: space-between; align-items: center; gap: 24px; }
.scroll-cue { display: flex; gap: 14px; align-items: center; text-decoration: none; font-size: .7rem; color: #d4dad5; }
.scroll-line { height: 30px; width: 1px; background: #d4dad577; }
.hero-toggle { display: inline-flex; align-items: center; gap: 9px; min-height: 44px; border: 1px solid #d2dbd144; border-radius: 100px; background: #18201b55; color: #e1e8df; padding: 8px 16px; font-size: .66rem; cursor: pointer; }
.hero-toggle:hover { background: #18201ba0; }
.hero-toggle [data-toggle-icon] { width: 12px; font-size: 10px; text-align: center; }
.section-nav { position: sticky; top: 0; z-index: 5; background: #ffffffef; border-bottom: 1px solid var(--line); backdrop-filter: blur(14px); }
.section-nav-inner { width: min(var(--width), calc(100% - 96px)); margin-inline: auto; display: flex; justify-content: space-between; gap: 32px; min-height: 68px; align-items: center; }
.section-brand { font-size: .63rem; font-weight: 600; letter-spacing: .13em; text-decoration: none; white-space: nowrap; display: inline-flex; align-items: center; gap: 12px; }
.brand-dot { display: inline-block; background: var(--accent); height: 6px; width: 6px; border-radius: 50%; flex-shrink: 0; }
.section-links { display: flex; gap: 28px; align-items: stretch; }
.section-links a { display: flex; align-items: center; min-height: 68px; font-size: .74rem; color: var(--muted); text-decoration: none; white-space: nowrap; border-bottom: 2px solid transparent; padding-top: 2px; }
.section-links a:hover, .section-links a[aria-current="location"] { border-bottom-color: var(--accent); color: var(--ink); }
.intro { padding-block: 108px 94px; }
.intro h2 { max-width: 900px; }
.intro-copy { font-size: 1.1rem; line-height: 1.85; max-width: 775px; color: var(--muted); margin-top: 28px; }
.research-facts { display: grid; grid-template-columns: 1fr 1.2fr 1fr; gap: 24px; padding-block: 30px; margin-top: 32px; border-top: 1px solid var(--line); border-bottom: 1px solid var(--line); }
.research-facts > div { display: flex; align-items: center; gap: 15px; font-size: .79rem; }
.fact-number { color: #9a7b12; font-size: .62rem; font-variant-numeric: tabular-nums; }
.abstract { max-width: 800px; margin-top: 28px; }
.abstract summary { display: inline-flex; gap: 36px; align-items: center; cursor: pointer; list-style: none; font-size: .8rem; }
.abstract summary::-webkit-details-marker { display: none; }
.abstract summary span { font-size: 1.2rem; line-height: 1; }
.abstract[open] summary span { transform: rotate(45deg); }
.abstract p { color: var(--muted); margin-top: 22px; font-size: .92rem; line-height: 1.85; }
.section { padding-block: 94px; }
#experiments { border-top: 1px solid var(--line); padding-top: 80px; }
.section-heading { margin-bottom: 64px; }
.section-heading > p:last-child { max-width: 710px; margin-top: 24px; color: var(--muted); font-size: 1rem; line-height: 1.85; }
.section-band { background: var(--wash); border-block: 1px solid #e9ece6; }
.experiment { margin-top: 64px; scroll-margin-top: 102px; }
.experiment + .experiment { margin-top: 80px; padding-top: 48px; border-top: 1px solid var(--line); }
.experiment-heading { display: grid; grid-template-columns: 1fr 1fr; gap: 50px; align-items: end; margin-bottom: 24px; }
.experiment-index { font-size: .6rem; color: #867330; letter-spacing: .13em; margin-bottom: 12px; font-weight: 600; }
.experiment h3 { font-size: 2.05rem; }
.experiment-subtitle { color: var(--muted); font-size: .83rem; margin: 10px 0 0; }
.experiment-description { color: var(--muted); font-size: .9rem; line-height: 1.8; margin: 0 0 1px; max-width: 500px; }
figure { margin: 0; min-width: 0; }
.video-shell { position: relative; aspect-ratio: 16 / 9; background: #e6e8e3; border-radius: 8px; overflow: hidden; }
.video-shell video { display: block; width: 100%; height: 100%; object-fit: contain; background: #e6e8e3; }
.experiment-figure figcaption { display: flex; gap: 24px; justify-content: space-between; margin-top: 14px; font-size: .65rem; color: var(--muted); letter-spacing: .01em; }
.duration { white-space: nowrap; font-variant-numeric: tabular-nums; color: #7d8680; }
.media-error, .noscript-media { position: absolute; inset: 0; margin: 0; padding: 32px; background: #ecefe8; display: flex; justify-content: center; align-items: center; gap: 16px; flex-direction: column; color: var(--ink); text-align: center; font-size: .9rem; }
.media-error a, .noscript-media a { display: inline-flex; align-items: center; gap: 8px; }
.section-note, .figure-note { font-size: .79rem; color: var(--muted); max-width: 820px; margin-top: 28px; }
.method-step { display: grid; grid-template-columns: 52px 1fr; gap: 20px; max-width: 820px; margin-block: 50px 26px; }
.step-number { padding-top: 2px; font-size: .72rem; color: #8d7622; line-height: 30px; border-top: 2px solid var(--accent); }
.method-step h3 { font-size: 1.55rem; letter-spacing: -.025em; }
.method-step p { color: var(--muted); font-size: .94rem; line-height: 1.85; margin: 14px 0 0; }
.paper-figure { margin-block: 30px 46px; }
.paper-figure > a { display: block; padding: 24px; background: #fff; border: 1px solid var(--line); border-radius: 6px; }
.paper-figure img { display: block; width: 100%; height: auto; }
.paper-figure figcaption { display: flex; flex-wrap: wrap; justify-content: space-between; gap: 10px 24px; margin-top: 12px; color: var(--muted); font-size: .66rem; }
.paper-figure figcaption span { display: inline-flex; align-items: center; gap: 6px; }
.paper-figure figcaption svg { width: 14px; height: 14px; }
.system-strip { border-top: 1px solid var(--line); padding-top: 25px; margin-top: 38px; }
.system-strip > span { font-size: .65rem; text-transform: uppercase; letter-spacing: .13em; color: var(--muted); }
.system-strip p { font-size: .78rem; margin: 10px 0 0; }
.system-strip b { color: #9da39a; padding-inline: 12px; font-weight: 400; }
.table-scroll { overflow-x: auto; border: 1px solid var(--line); border-radius: 6px; margin-top: 38px; }
table { border-collapse: collapse; width: 100%; min-width: 680px; font-size: .86rem; }
caption { text-align: left; padding: 19px 24px; font-size: .75rem; background: #f7f8f5; }
caption span { float: right; font-size: .65rem; color: var(--muted); }
th, td { padding: 18px 24px; border-top: 1px solid var(--line); text-align: center; white-space: nowrap; font-variant-numeric: tabular-nums; }
th { font-weight: 500; }
th[scope="row"], th[rowspan] { text-align: left; }
thead th { font-size: .76rem; }
.ours { background: #f8f4df; }
.result-highlight { font-weight: 600; color: #625110; }
.table-note { font-size: .75rem; color: var(--muted); margin-top: 17px; }
.table-note strong { font-weight: 500; color: var(--ink); }
.morphology-heading { margin-top: 75px; max-width: 780px; }
.morphology-heading h3 { font-size: 1.6rem; }
.morphology-heading p { margin-top: 18px; color: var(--muted); font-size: .95rem; }
.results .paper-figure { margin-bottom: 14px; }
.figure-note { margin-top: 14px; font-size: .72rem; }
.limitations { padding-block: 60px 100px; border-top: 1px solid var(--line); }
.limitations-layout { display: grid; grid-template-columns: 1fr 1.2fr; gap: 70px; }
.limitations h2 { font-size: 2.3rem; }
.limitations-layout p { color: var(--muted); font-size: .86rem; line-height: 1.85; }
pre { padding: 24px; border: 1px solid var(--line); background: var(--wash); overflow-x: auto; border-radius: 6px; font-size: .82rem; }
.footer { display: flex; align-items: center; gap: 30px; padding-block: 32px 42px; border-top: 1px solid var(--line); font-size: .69rem; color: var(--muted); }
.footer a { display: inline-flex; align-items: center; gap: 10px; text-decoration: none; }
.footer a:hover { color: var(--ink); }
.footer-title { margin-right: auto; color: var(--ink); }
.footer svg { width: 17px; height: 17px; }
@media (min-width: 1600px) { .hero-content { padding-top: 13vh; } }
@media (max-width: 1024px) {
  .hero { min-height: max(800px, 100svh); }
  .hero-nav, .hero-content, .hero-bottom { width: calc(100% - 72px); }
  .hero-content { padding-top: 100px; }
  .hero-nav-links { gap: 25px; }
  .hero-shade { background: linear-gradient(90deg, #111a17dc, #111a1780 65%, #111a1733), linear-gradient(0deg, #111a1777, transparent 30%); }
  .container, .section-nav-inner { width: calc(100% - 64px); }
  .section-links { gap: 20px; }
  .section-brand { font-size: .56rem; letter-spacing: .08em; gap: 8px; }
  .research-facts { grid-template-columns: 1fr; gap: 16px; }
  .experiment-heading { gap: 30px; grid-template-columns: 1fr 1.2fr; }
  .experiment h3 { font-size: 1.8rem; }
  .limitations-layout { gap: 40px; }
}
@media (max-width: 600px) {
  body { font-size: 16px; }
  html { scroll-padding-top: 80px; }
  .container { width: calc(100% - 40px); }
  .hero { min-height: max(740px, 100svh); }
  .hero-nav, .hero-content, .hero-bottom { width: calc(100% - 40px); }
  .hero-nav { padding-top: 26px; }
  .hero-nav-links { gap: 17px; }
  .hero-nav-links a { font-size: .68rem; }
  .hero-nav-links a:last-child { display: none; }
  .wordmark span { font-size: 8px; }
  .wordmark svg { width: 29px; height: 29px; }
  .hero-content { padding-top: 111px; padding-bottom: 125px; }
  h1 { font-size: clamp(3.25rem, 12.7vw, 4.5rem); line-height: 1.045; }
  .hero-eyebrow { font-size: .54rem; letter-spacing: .12em; margin-bottom: 22px; }
  .hero-summary { font-size: .88rem; max-width: 310px; margin-top: 26px; }
  .paper-authors { margin-top: 24px; font-size: .84rem; }
  .author-list { gap: 8px 18px; }
  .author-affiliations { font-size: .71rem; }
  .author-notes { font-size: .65rem; }
  .hero-actions { gap: 10px; margin-top: 28px; }
  .button { min-height: 47px; gap: 13px; padding: 10px 14px; font-size: .7rem; }
  .hero-media img, .hero-media video { object-position: 61% center; }
  .hero-shade { background: linear-gradient(90deg, #111a17dd, #111a17a0 70%, #111a1770), linear-gradient(0deg, #111a1780, transparent 35%); }
  .hero-bottom { bottom: 22px; gap: 16px; }
  .scroll-cue { font-size: .59rem; gap: 9px; }
  .hero-toggle { padding: 8px 12px; font-size: .6rem; }
  .section-nav-inner { width: 100%; min-height: 58px; }
  .section-brand { display: none; }
  .section-links { justify-content: space-between; gap: 22px; overflow-x: auto; width: 100%; padding-inline: 20px; }
  .section-links a { min-height: 58px; font-size: .68rem; }
  .intro { padding-block: 72px 60px; }
  .eyebrow { font-size: .62rem; margin-bottom: 20px; }
  h2 { font-size: 2.1rem; line-height: 1.15; }
  .intro-copy { font-size: .96rem; margin-top: 24px; }
  .research-facts { padding-block: 25px; margin-top: 25px; }
  .research-facts > div { font-size: .77rem; }
  .abstract summary { font-size: .76rem; }
  .abstract p { font-size: .86rem; }
  .section { padding-block: 64px; }
  #experiments { padding-top: 60px; }
  .section-heading { margin-bottom: 40px; }
  .section-heading > p:last-child { font-size: .92rem; margin-top: 22px; }
  .experiment { margin-top: 40px; }
  .experiment + .experiment { margin-top: 44px; padding-top: 35px; }
  .experiment-heading { display: block; margin-bottom: 20px; }
  .experiment h3 { font-size: 1.85rem; }
  .experiment-index { font-size: .56rem; margin-bottom: 10px; }
  .experiment-subtitle { font-size: .78rem; margin-top: 8px; }
  .experiment-description { font-size: .84rem; margin-top: 17px; }
  .video-shell { border-radius: 5px; }
  .experiment-figure figcaption { font-size: .59rem; line-height: 1.7; gap: 14px; margin-top: 11px; }
  .section-note { font-size: .75rem; margin-top: 24px; }
  .method-step { grid-template-columns: 30px 1fr; gap: 16px; margin-top: 38px; }
  .method-step h3 { font-size: 1.36rem; }
  .method-step p { font-size: .85rem; margin-top: 12px; }
  .step-number { font-size: .66rem; }
  .paper-figure { margin-block: 24px 38px; }
  .paper-figure > a { padding: 12px 8px; }
  .paper-figure figcaption { font-size: .6rem; }
  .system-strip p { font-size: .71rem; line-height: 2; }
  .system-strip b { padding-inline: 6px; }
  .table-note { font-size: .72rem; }
  .morphology-heading { margin-top: 52px; }
  .morphology-heading h3 { font-size: 1.4rem; }
  .morphology-heading p { font-size: .85rem; }
  .figure-note { font-size: .68rem; }
  .limitations { padding-block: 45px 60px; }
  .limitations-layout { display: block; }
  .limitations h2 { font-size: 2rem; margin-bottom: 26px; }
  .limitations-layout p { font-size: .84rem; }
  .footer { flex-wrap: wrap; gap: 20px; padding-block: 26px 35px; font-size: .64rem; }
  .footer-title { width: 100%; }
  .back-top { margin-left: auto; }
}
@media (max-width: 360px) {
  .hero-actions { gap: 8px; }
  .button { gap: 9px; padding-inline: 12px; }
  .button-note { display: none; }
  .hero-summary { font-size: .8rem; }
  .scroll-cue { max-width: 160px; line-height: 1.5; }
}
@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { transition: none !important; }
}